Suddenly, Captain Dylan Hunt isn't himself.

The Andromeda crew is traversing deep space, navigating the ship through a maze of space anomalies and wormholes, when a surprise attack by an unidentified fleet of ships forces them into a space anomaly. The Andromeda crosses into another dimension, where their captain is a green-skinned, cold, logical, Nietzschean Dylan who's more reptilian than human. The crew must band together and choose between obedience and mutiny as they follow a dangerous course into an unknown dimension of space with an alien captain they cannot trust.

Meanwhile, the real Dylan commands a very different Andromeda. His new crew of green-skinned reptilians is engaged in a deadly conflict with rebels who, in Dylan's original dimension, are patriots of the civilization he has tried to restore. His new crew seems to be involved in a vast interstellar conquest that he must stop, and he realizes he must defuse a potentially explosive conflict if he's to survive his alien ship.

The decisions both Andromeda crews and captains make may cause ripple effects that could save or destroy both dimensions.