Is this journey's end . . . or just the beginning?

Toklo, Kallik, Lusa, and Ujurak have finally reached the Last Great Wilderness, the legendary bear paradise they've been searching for. But while his companions think they've come to the end of their long journey, Ujurak feels a deep unrest. Is this truly where they're meant to be?

In the Last Great Wilderness, one by one the bears begin to remember their true natures. Toklo feels the urge to hunt caribou and mark his territory as a brown bear should, and Kallik feels the pull of the ice within her. It's only Lusa, happy just to be in the wild, who fears the day when her friends will leave her to follow their own paths.

As the bears adjust to this new life, disaster strikes. The friends are forced to venture into the world of the flat-faces to save the life of one of their own. Once there, the end of their journey seems farther away than ever, as a new path spreads out before them.